SCLERANTHUS VERTICILLATUS
Common Names:- Annual knawel
Homotypic Synonyms:- Scleranthus annuus subsp. verticillatus
Meaning:- Scleranthus (Gr) With hard thorns or spines.
Verticillatus (L) Having whorls (several leaves or flowers all arising at
the same level on the stem).
General description:- Small annual or occasionally biennial herb.
Stems:-
1) 5-15 cm.
Leaves:-
1) 5-10 mm linear, channelled, obtuse, ciliate at the base.
Flower:-
1) Subsessile in terminal and auxillary clusters.
2) Sepals, 1-1.5 mm. unequal, incurved, equalling or longer than the perigynous
zone, acute, almost without membranous margins.
3) Petals, lacking.
Fruit:-
1) Nutlet, 1·2-5·5 mm, indehiscent, 1(2)-seeded, enclosed by the hardened wall of
the perigynous zone and the persistent sepals, which are shed with it.
2) Seeds, lenticular, smooth.
Key features:-
1) Sepals, usually 1-1.5 mm, unequal, incurved, almost without membranous
margins.
Habitat:- Rocky and gravelly slopes, roadsides, open coniferous woodland. 0-1200
m. (-1700 m. in dolines).
Distribution:- Mainland Greece, Peloponnisos, Kefallinia. - Widespread in S & C
Europe and the Mediterranean region eastwards to Syria. Rare on Crete, scattered
across the four main massifs
Flowering time:- Apr-June.
Photos by:- Courtesy of Wiki-Commons.